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(Click for frequent words.) 54 National Marine Sanctuaries 54 marine sanctuaries 53 coral reef 51 Northwestern Hawaiian Islands 49 Estuarine Reserve 49 Stellwagen Bank National 48 Las Baulas 48 kelp forest 47 Coiba 47 Humpback Whale National 47 Farallones 46 Looe Key 46 Maritime Museum 46 Sanibel Captiva Conservation 46 Marine Aquarium 46 Marine Protected Areas 46 reef 46 Dry Tortugas 45 Artificial Reef Program 45 coral 45 Papahanaumokuakea Marine National 45 Aquatic Preserve 45 marine ecosystem 45 reefs 45 Coral Reef Conservation 45 Haystack Rock 45 Papahānaumokuākea Marine National 45 Coral Reef Ecosystem 45 coral reef ecosystem 44 Davidson Seamount 44 Healthy Beaches 44 National Seashore 44 National Estuarine Research 44 Hawaiian Islands Humpback Whale 44 Underwater Preserve 44 Data Buoy 44 Ahihi Kinau 44 Hawai'i Volcanoes National 44 marine mammal 44 NERR 44 Barataria Preserve 44 sea turtle 44 Tortuguero 44 intertidal 44 Bunaken 43 Belize Barrier Reef 43 Keawakapu 43 Ecological Reserve 43 marine mammal sanctuaries 43 Nihoa Island 43 Gwaii Haanas 43 Atmospheric Administration 43 Stellwagen Bank 43 Tubbataha 43 San Dieguito Lagoon 43 Estuary Program 43 Marine Mammal Laboratory 43 leatherback sea turtles 43 Ocean Observing 43 kelp beds 43 Rose Atoll 43 Laysan Island 43 NWHI 43 coral reefs 43 MBARI 43 nearshore waters 43 Marine Mammal Stranding Network 43 Waquoit Bay 42 Malama Maunalua 42 Egmont Key 42 Great Bear Rainforest 42 Estate Maho 42 Similan 42 longline fishery 42 Drakes Estero 42 Dungeness Spit 42 Vancouver Aquarium 42 Oceanographic 42 Quartermaster Harbor 42 tidepools 42 corals 42 elkhorn coral 42 eelgrass 42 Ningaloo 42 Monk Seal 41 Marine Protected 41 monk seals 41 Marine Mammals 41 Marine Mammal Response 41 Rookery Bay Reserve 41 Marine Mammal 41 Recreation Area 41 Hawaiian monk seals 41 rocky reefs 41 shoreline 41 marine mammals 41 barrier reef 41 sea otters 41 SeaDoc Society 41 Molokini 41 snowy plover 41 Hol Chan Marine 41 Fisheries 41 Kure Atoll 41 Regional Fishery Management 41 leatherback sea turtle 41 sea turtle conservation 41 seashore 41 Leatherback Turtle 40 WiLDCOAST 40 coral formations 40 French Frigate Shoals 40 Loggerhead 40 marine ecosystems 40 Bacalar Chico 40 Gwaii Haanas National 40 vulnerable marine ecosystems 40 coastal ecosystems 40 pawikan 40 NOAA 40 Estuarine 40 reefs teeming 40 seabeds 40 Sustainable Fisheries 40 Preserve 40 Estuaries 40 Fishery Management Council 40 Nautical Museum 40 Deep Submergence Facility 40 Tropical Botanical Garden 40 Elkhorn Slough 40 vaquita 40 seagrass meadows 40 Great Barrier Reef 40 Shellfish Sanitation Program 40 Farallon Islands 40 manta rays 40 seals sea otters 40 estuarine 40 salt marsh 40 Tsunami Hazard Mitigation 40 Marine Fisheries Service 40 Atmospheric Administration Fisheries 40 estuarine habitat 40 Lighthouse Preservation 40 sea otter 40 Albemarle Pamlico 40 eelgrass meadows 40 Farallons 40 Rookery Bay 40 Sean Van Sommeran 40 Skan Bay 40 Mealy Mountains 40 Seaside Aquarium 40 Laura Engleby 40 Pelagic Shark Research 40 pristine coral reef 40 seagrasses 40 spiny lobsters 40 Ka Iwi 40 salt marsh habitat 40 Mote Marine 40 Aquarium 40 Marine Mammal Center 39 groundfish stocks 39 Bolsa Chica Wetlands 39 saltwater marsh 39 Pamilacan 39 Morro Rock 39 sea turtles 39 Nauset Marsh 39 CoML 39 Tiffany Boothe 39 Farallones Marine 39 Stranding Network 39 Palmyra Atoll 39 VINP 39 Wildlife Federation NWF 39 Abrolhos 39 submersible Alvin 39 Shirley Pomponi 39 Haida Heritage Site 39 Birch Aquarium 39 Cape Lookout 39 Kayak Tours 39 oyster reef 39 Tubbataha Reef 39 artificial reef 39 dune lakes 39 Cape Perpetua 39 shrimp trawling 39 Randy Kosaki 39 elkhorn corals 39 Point Pelee National 39 biosphere reserve 39 marine turtles 39 marine 39 marine mammal strandings 39 whale 39 Point Fermin 39 Solitary Islands 39 Morro Strand 39 estuary 39 northwestern Hawaiian Islands 39 Harmful Algal Blooms 39 Tomales Bay 39 Bodie Island 39 Great Lakes Shipwreck 39 Sea otters 39 Dabob Bay 39 shallow reefs 39 Waikiki Aquarium 39 coral reef habitat 39 Hawaiian Archipelago 39 tortoise habitat 39 ocean ecosystems 39 Sperm whale 39 eel grass 39 Loggerhead Marinelife Center 39 Cay Sal Bank 39 marine biodiversity 39 gill nets 39 Año Nuevo 39 artist Wyland 38 CoastWatch 38 turtle nesting beaches 38 shorelines 38 coastal waters 38 striped bass fishery 38 Po'ipu Beach 38 Fur Seal 38 Bolinas Lagoon 38 endangered leatherback sea turtles 38 subtidal 38 Randi Sokoloff 38 oyster reefs 38 Reefs 38 Recreational Fisheries 38 Cosco Busan spill 38 Hawksbill Turtle 38 BP Deepwater Horizon 38 NOAA Fisheries Service 38 threatened marbled murrelet 38 Fort Matanzas 38 Queets River 38 Mote Marine Laboratory 38 tidal pools 38 Kopachuck 38 IODP 38 Atmospheric Administration Coral Reef 38 Tubbataha Reefs 38 inland waters 38 Midway Atoll 38 leatherback turtles 38 Ostego Bay 38 Bolsa Chica Conservancy 38 museum Te Papa 38 Dungeness River 38 Tatoosh Island 38 Earthquake Hazards Reduction 38 NOAA Fisheries 38 Matagorda Bay 38 L' Orca 38 endangered sea turtles 38 Marine Ecosystems 38 Nature Conservancy 38 Osa Peninsula 38 Foulweather Bluff 38 Matagorda Island 38 Deepwater Horizon Oil Spill 38 tidal wetlands 38 Tortugas Ecological Reserve 38 Sarasota Mote Marine 38 Clayoquot Sound 38 endangered leatherback turtles 38 Ocean Acidification Program 38 cod fisheries 38 salt marshes 38 Dolphin Encounters 38 monk seal 38 OSPR 38 fairy tern 38 Gahirmatha Marine Sanctuary 38 nesting seabirds 38 NOAA Coral Reef 38 Akaroa Harbour 38 La Jolla Underwater 38 Hawaiian monk seal 38 Cape Flattery 38 Blueways 38 Tarutao 38 Wilderness Preservation 38 Schoodic Point 38 Artificial Reefs 38 Estuaries Program 38 Cobscook Bay 38 sealife 38 Oceanography Centre 38 Wetland 38 fragile marine ecosystems 38 coastal bays 38 Basking Shark 38 Madrona Marsh 38 Coastal Bays 38 seagrass beds 38 Coastkeeper 38 coral ecosystems 38 seagrass 38 Florida Oceanographic Coastal 38 oystercatcher 38 Kasatochi 38 Ka'ena Point 38 Marinelife Center 37 Beidler Forest 37 John Pennekamp Coral 37 Piedras Blancas 37 plover habitat 37 Marine Fisheries 37 pristine coral reefs 37 Maro Reef 37 Cape Hatteras lighthouse 37 Cape Arago 37 kelp 37 mussel beds 37 Lad Akins 37 Beaufort Inlet 37 Alaska Inside Passage 37 navigational buoys 37 Peril Strait 37 Sea Turtles 37 orca whale 37 scallop dredging 37 Hawaiian Islands 37 Point Ano Nuevo 37 Quirimbas 37 shellfish beds 37 National Marine Fisheries 37 Pasagshak 37 Carol Diebel 37 marine habitats 37 seabed 37 Yukon Charley 37 Artificial Reef 37 Exxon Valdez Oil Spill 37 dolphins sea lions 37 Elk Refuge 37 Napeague Harbor 37 oyster beds 37 Lophelia II 37 marine mammal rescues 37 Russell Reichelt 37 Oceanography UC 37 inter tidal 37 halibut fishing 37 Oil Spill Response 37 leatherback turtle 37 Peconic Estuary 37 SEA LIFE Aquarium 37 Santa Monica BayKeeper 37 Point Pinole 37 Point Buchon 37 coral reef ecosystems 37 Robson Bight 37 Hubbs SeaWorld 37 shellfish aquaculture 37 Lenfest Ocean 37 turtle nests 37 Paulo Vecina 37 Undersea Research 37 Offshore Aquaculture 37 Mote Aquarium 37 Fishery Management 37 Alameda whipsnake 37 Pemaquid Point 37 mollusks 37 Vermilion Cliffs 37 spokeswoman Alexia Retallack 37 tern colony 37 marine biologists 37 Oscar Elton Sette 37 Moonstone Beach 37 loggerhead sea turtles 37 Wildcoast 37 kelp forests 37 Rimpac 37 mudflat 37 Maui dolphin 37 Kauai Na Pali 37 Atmospheric Administration NOAA Fisheries 37 gillnets 37 Ecological Analysis 37 Guana Reserve 37 Pew Oceans Commission 37 intertidal zone 37 snowy plovers 37 Coastal Ecosystems 37 Delaware Inland Bays 37 Wildlife Federation 37 Bottlenose Dolphins 37 Tobay Beach 37 Audubon Society 37 Coupeville Wharf 37 prawn trawling 37 Guadalupe Dunes 37 estuaries 37 Yaquina Head 37 dolphins frolic 37 Cape Cod Stranding Network 37 jumbo squid 37 USS Monitor 37 finfish 37 abalone 37 Punta Nizuc 37 Lightstation 37 fisheries 37 Petit Manan 37 diamondback terrapin 37 Papahanaumokuakea 37 whales 37 sea lion 37 Allens Pond 37 Kemp ridley 37 clam harvesting 37 Malibu Lagoon 37 inshore artificial reefs 37 Cabo Pulmo 37 Loggerhead Sea 37 loggerhead sea turtle 37 Coastweeks 37 eelgrass beds 37 Guana Tolomato Matanzas 37 lobster fishery 37 DOCARE 36 Caxambas Pass 36 Hubbard Glacier 36 Ventana Wilderness Society 36 gray whale 36 Ramsar wetland 36 gillnet 36 oyster restoration 36 Nomans Land 36 Willapa Bay 36 Co op Chemists 36 green hawksbill 36 Response OSPR 36 Wildlife Refuge Complex 36 pier pilings 36 Drakes Bay 36 Dosewallips 36 Hydrographic 36 Barataria Basin 36 Maunalua Bay 36 longlines 36 groundfish fisheries 36 Lophelia 36 loggerhead turtle 36 abandoned crab traps 36 Packery Channel 36 Apo Reef 36 Leatherback turtles 36 Fort Cronkhite 36 Haleakala 36 Ataxia Foundation 36 La Parguera 36 Atlantic Bottlenose dolphins 36 Cape Kiwanda 36 recreational fishery 36 mussel harvesting 36 seabird nesting 36 Cape Poge 36 Estuarine Research Reserve 36 Risso dolphins 36 crab fishery 36 Recreational Area 36 tidewater goby 36 Mako shark 36 ocean 36 eared grebe 36 Phong Nha Ke Bang 36 Cape Meares 36 Hector dolphins 36 bioluminescent bay 36 tidal wetland 36 Sulu Sulawesi 36 Tonquin Foundation 36 Coastal Wetlands 36 derelict crab pots 36 Mass Audubon Wellfleet 36 Underwater Archaeology Branch 36 Marblehead Lighthouse 36 Steller sea lion 36 Oceanography 36 Galapagos sharks 36 Dyes Inlet 36 Sealife 36 Risso dolphin 36 Monomoy Island 36 sea turtle populations 36 cove 36 Salish Sea 36 Manatee Pocket 36 Vernal Pool 36 Coral Reef Ecosystems 36 endangered manatees 36 wildlife refuges 36 shallow lagoon 36 Park Authority GBRMPA 36 Fakahatchee Strand 36 Kimbe Bay 36 coral habitats 36 senior aquarist 36 Campland 36 Habitat Stewardship 36 cetaceans whales dolphins 36 oiled wildlife 36 coral colonies 36 whales beached themselves 36 Cocos Island 36 NOAA marine mammal 36 seafloor 36 longline fishing 36 diver swims 36 Lituya Bay 36 coral fringed 36 endangered whales 36 Frontier Trails 36 endangered seabird 36 sea turtle species 36 marbled murrelets 36 Vanishing Treasures 36 Florida scrub jay 36 visit www.oceanconservancy.org 36 Shinnecock Inlet 36 nearshore reefs 36 rocky cove 36 Cook Inlet Keeper 36 seabird 36 Aquatic Invasive Species Act 36 Cuvier beaked whale 36 Cape Disappointment 36 cabled observatory 36 Auke Bay Laboratory 36 migrating whales 36 coastal estuaries 36 NOAA fisheries 36 Maritime Archaeology 36 shorebird habitat 36 Aquarium Seafood 36 squid 36 Mokulua Islands 36 dolphins 36 geoduck farming 36 tar mats 36 scallop fishery 36 crab trap 36 Kuhio Beach 36 Izembek 36 Muller Karger 36 Heritage Canalway 36 oil spill 36 aquarium 36 fringing reefs 36 shellfish harvesting 36 Point Piedras Blancas 36 Blowing Rocks Preserve 36 Jurassic Coast 36 Willapa 36 Chagres River 36 sea turtles nest 36 Lt. Todd Vorenkamp 36 Kahoolawe Island 36 Playalinda Beach 36 Snorkelers 36 porpoises 36 Visitor Center 36 reef ecosystems 36 Kualoa 35 Anacapa Island 35 Coral reef 35 submerged aquatic vegetation 35 APVA Preservation Virginia 35 Atmospheric Administration fisheries 35 estuarine habitats 35 hawksbill turtle 35 Laguna Madre 35 whales seabirds 35 Bill Hanshumaker 35 Marine Mammal Protection 35 Vessel Tracking 35 seabirds 35 dolphins whales 35 marine biologist 35 Conservancy 35 tidal salt marsh 35 GGNRA 35 whale watching 35 saltwater lagoon 35 David Wachenfeld 35 Agate Fossil Beds 35 Santa Barbara Channelkeeper 35 Jiggs Landing 35 sea turtles grouper 35 Marine Mammal Rescue 35 underwater archeology 35 National Oceanic 35 Whale Watch 35 coastal wetlands 35 equatorial waters 35 intertidal zones 35 fishery 35 Norway Svalbard 35 Cape Sable 35 Studland Bay 35 Nissequogue River 35 harmful algae blooms 35 Kenai Fjords National 35 wolffish 35 sea creatures 35 Bolsa Chica 35 vessel groundings 35 seamounts 35 Makah tribe 35 Boat Shows 35 shipwrecked Spanish galleon 35 Beluga whales 35 fragile reef ecosystems 35 turtles nesting 35 Wildlife Refuge 35 Farne Islands 35 humpback whales migrate 35 Golfo Dulce 35 nudibranch 35 breakwater 35 sea turtle nest 35 Nassau grouper 35 harlequin duck 35 fragile marshes 35 artificial reefs 35 killer whale 35 Maui dolphins 35 abalones 35 Wyoming Grand Teton 35 tidal tributaries 35 oarfish 35 Ao Nuevo 35 Cu Lao Cham 35 Tigertail Beach 35 vernal pool 35 Copano Bay 35 Geologic Trail 35 Puget Sound orcas 35 horseshoe crab spawning 35 reef habitats 35 Oil Spill Prevention 35 Roosevelt Inlet 35 Cruise Ship Terminal 35 Laguna Atascosa National 35 Caretta caretta 35 elkhorn 35 Vessel Monitoring 35 olive ridley 35 sheltered bays 35 nesting sea turtles 35 rough toothed 35 SRIA 35 Cockroach Bay 35 manatee 35 Virginia Aquarium Stranding 35 Kouchibouguac National 35 shorebird 35 Grays Harbor Historical 35 Historic Blakeley 35 Cypress Slough 35 Cape Hatteras NC 35 sunken vessel 35 biotoxin monitoring 35 belugas 35 marine ecology 35 Exxon Valdez ran aground 35 biological oceanography 35 Federal Subsistence 35 Dingmans Falls 35 Humpback whale 35 Ogasawara Islands 35 Masonboro Island 35 fringing reef 35 Stearns Wharf 35 FWC 35 Magnuson Stevens Fishery Conservation 35 Clean Diesel Campaign 35 Kilo Moana 35 UMCES 35 Oceans Atmosphere Fisheries 35 Dall porpoise 35 estuarine ecology 35 Unalaska Island 35 nesting colony 35 Odyssey Explorer 35 Neil Cramond 35 FWRI 35 Saugatuck Dunes 35 Refuge Visitor 35 Jacob Verhoef 35 Delnor Wiggins 35 corals sponges 35 Artificial reefs 35 whale sanctuary 35 Visayan Sea 35 diamondback terrapins 35 Schoodic 35 UDT SEAL 35 Inland Waterway 35 freshwater wetland 35 Beluga Whales 35 Johnstone Strait 35 Teshekpuk Lake 35 haulout 35 Cape Hatteras 35 Cape Romain 35 rocky intertidal 35 hawksbill turtles 35 Salem Sound Coastwatch 35 San Dieguito River 35 Ocean Conservancy 35 IOOS 35 JOIDES Resolution 35 Sperm Whale 35 starfish sea urchins 35 shorebird nesting 35 Jane Lubchenko 35 Garvies Point Museum 35 OSV BOLD 35 otters 35 inshore waters 35 Pulau Weh 35 Honokohau Harbor 35 O'ahu Kaua'i 35 turtle nesting sites 35 ship groundings 35 lighthouse 35 Moapa dace 35 Waquoit Bay National 35 leatherback 35 La Jolla cove 35 Duties Lazing around 35 inshore fisheries 35 Muskeget Island 35 basking shark 35 Oceanographic Society 35 upland habitats 35 oceanographic 35 Brad Pettinger 35 Los Penasquitos Canyon 35 dolphins porpoises 35 St. Lucie Inlet 35 Blanding turtle 35 nesting turtles 35 River Estuary 35 Killer Whale 35 dugongs 34 endangered manatee 34 Environmental Laboratory Accreditation 34 shrimp fishery 34 groundfish 34 Pu'uhonua o Honaunau 34 Motuihe 34 Greg Siekaniec 34 Scuba dive 34 USGS 34 Hector dolphin 34 Shellfish Sanitation 34 oyster dredge 34 West Quoddy Head 34 Arctic Ocean 34 recreational fishers 34 manatee carcasses 34 coastal headlands 34 Hawksbill turtle 34 shoaling 34 Scripps Oceanography 34 Laura Engleby biologist 34 Sespe Wilderness 34 Whales dolphins 34 Similan Islands 34 Bering seas 34 tidal marsh 34 Atmospheric Administration NOAA 34 Western Snowy Plover 34 Bahia Honda 34 Oyster Restoration 34 harbor porpoise 34 Tarkine wilderness 34 Kristin Laidre 34 Marine Mammal Stranding 34 Galápagos 34 vernal pool species 34 Porpoise Point 34 swordfish fishery 34 rookery 34 Floras Lake 34 Great Sand Dunes 34 Savannas Preserve State 34 oiled sea turtles 34 Foundation NSF 34 volcanic archipelago 34 Deepwater Horizon spill 34 Keewaydin Island 34 Coast Gaurd 34 Polar Programs 34 Salt Ponds 34 stony coral 34 Ningaloo Marine 34 Kaimana Beach 34 endangered leatherback 34 Chen Lujun 34 humpback whales 34 maritime archaeologist 34 Cook Inletkeeper 34 Molokini Crater 34 George Durgerian 34 Cape Otway 34 leatherback nesting 34 aquatic species 34 sandy cove 34 WHOI 34 snorkelers 34 freshwater lakes 34 Cape Lookout Lighthouse 34 riparian habitats 34 basking sharks 34 Shedd 34 lightship 34 inland bays 34 DLNR 34 Mote Marine Laboratories 34 orca whales 34 Least Terns 34 Eelgrass 34 tropical coral reefs 34 Whydah 34 Body washes ashore 34 Ano Nuevo 34 Sengekontacket Pond 34 Wildlife Preserve 34 Gaviota Coast 34 GCRL 34 dolphin 34 renourished 34 abalone fishery 34 whale disentanglement 34 seabird breeding 34 Kaho'olawe 34 Haro Strait 34 marine ecologist 34 Shark Savers 34 Assessment Governing 34 Wakatobi 34 Hopewell Rocks 34 Fishing Guides 34 nesting beaches 34 Tumon Bay 34 Atmospheric Administration Emergency Response 34 shipwreck artifacts 34 Santa Barbara Botanic Garden 34 Harbor Branch Oceanographic 34 sperm whales 34 common murres 34 mangrove ecosystems 34 Great Lakes Ecosystem 34 Coral Reefs 34 Dalco Passage 34 oyster aquaculture 34 Boca Grande Lighthouse 34 Clean Diesel Funding 34 Bangtao 34 tuna fishery 34 Makushin Bay 34 Wassaw Island 34 surf clams 34 River Lagoon 34 Wailua River 34 Padre Balli Park 34 bald eagle nest 34 Seashore 34 Overmountain Victory 34 Recreational fishermen 34 nonregulatory 34 humpback whale 34 Seagrass 34 Gray Whales 34 northern fulmars 34 red grouper 34 SEE Turtles 34 whales sea lions 34 Kewalo Basin 34 Spurn Point 34 greater shearwaters 34 Humboldt squid 34 mangrove islands 34 BICREF 34 Leatherback sea turtles 34 Tomales Point 34 Kejimkujik National 34 Charterboat Association 34 geoduck 34 Skomer 34 Sandy Hook NJ 34 Duxbury Reef 34 endangered Hawaiian monk 34 Bill Coklough 34 shellfisheries 34 Twanoh 34 whale shark 34 Lubec Maine 34 Hokule'a 34 Blair Mase Guthrie 34 Audobon Society 34 Ramsar wetlands 34 Caspersen Beach 34 Tiritiri Matangi 34 Tuckerton Seaport 34 tideland 34 Administrator Jane Lubchenco 34 Skomer Island 34 Oil Spill 34 vaquita porpoise 34 seal rookery 34 oceans coasts 34 coral cays 34 Sandy Neck 34 spiny lobster 34 UCSD Scripps Institution 34 JD Murphree 34 coral Enzler 34 Musandam peninsula 34 mooring buoys 34 nosed dolphin 34 Mo'omomi 34 Na Pali Coast 34 eel fishery 34 Makapuu 34 Beluga Whale 34 coastal dune 34 rhinoceros auklets 34 Superintendent Dave Fronzuto 34 Chenier Plain 34 Tybee Island Marine 34 dugong 34 tidal marshes 34 Stingray Feeding Program 34 Auyuittuq National 34 Kemp ridley turtle 34 Lahaina Harbor 34 estero 34 Bottom trawling 34 Whaleback Lighthouse 34 smalltooth sawfish 34 sea turtle nesting 34 oyster larvae 34 Bob DeGross 34 aquatic habitats 34 Lower Otay Reservoir 34 Presidio Trust 34 zoo giant panda 34 Grayton Beach 34 Coral Triangle 34 mangrove forests 34 Svalbard Islands 34 compulsory pilotage 34 Unimak Island 34 Fishing Pier 34 aquatic fauna 34 turtles 34 Artificial Reef Society 34 Fiordland 34 Coast Guard 34 Response Agency NOSDRA 34 nautical charting 34 bottomfish 34 shallow waters 34 National Oceanic Atmospheric 34 horseshoe crabs 34 turtle nesting 34 Fort Wetherill 34 sharks sea turtles 34 pristine coastline 34 seal pups 34 ACND 34 KP2 34 Whale Watching 34 tiger salamanders 34 Arctic Coring Expedition 33 Coral Reef 33 Porpoise 33 harmful algal blooms 33 Maryland Coastal Bays 33 Vertebrate Zoology 33 shallower reefs 33 shellfish