monoclinic diatoms bacteriochlorophyll chlorophyll hemes tropane cytochrome Rhodophyta carotenes subleq Archaeplastida YFP algae heterokont anucleate unicellular phe Chlamydomonas dinoflagellates carotenoids phycobiliproteins 4r filamentous flagellate Titanosauridae mitosomes haptophytes pigments thylakoids plastid azo Proterospongia