ornithopod supercellular oviraptorids digitigrade Heterodontosaurus Chirostenotes tyrannosaurids Orodromeus Hydantoin bipedality Alioramus azhdarchoid tahrs piscivore lambeosaurines Euparkeria theropod hooved Nemegtosaurus Khillari isc intergrading cavies animal omomyids Kerodon Galeus sault hindlimbs Bactrosaurus infall pterosaurs parvus intragenomic slipface Asperula relative pteryx Pakicetus